When Charlotte Lucas's twin brother returns to Meryton after four years in the navy, he rivals even Mrs. Bennet as a meddling matchmaker. Captain Oliver Lucas introduces his two dearest friends, Elizabeth Bennet and Captain Fitzwilliam Darcy, and their discreet courtship begins on the very same night that Elizabeth discovers her childhood friend harbors a secret which will later alter the course of their lives.

As Elizabeth's romance blooms, Jane faces an ultimatum from her mother, and takes drastic measures to secure her engagement to Mr. Bingley. Jane's collusion with Caroline Bingley estranges her from her sisters when she causes a calamitous chain reaction that jeopardizes the happiness of more than one of her sisters, and has even more disastrous results for the Darcy family.

In the two years that Elizabeth and Captain Darcy are separated, their lives are consumed by tragedy and despair, though neither can let go of their memories of the love they once shared. Meanwhile, the Bingley marriage flounders when both Jane and Charles Bingley are haunted by the mistakes of their past, led afoul by a cruel and conniving Caroline.

When Elizabeth　Captain Darcy are finally reunited, they must confront their painful history before they can begin anew together, and in doing so give hope to those they love. The long-suffering Bingleys attempt their own reconciliation with help from an older and wiser Lydia Bennet, as well as the charming and mischievous Colonel Fitzwilliam. A house party full of familiar faces and new characters alike sets the scene for romance, and amidst the revelry and reminiscing, more than one couple finds their happily ever after, proving that time does heal all wounds.