After being in four weddings in four years, Donna has a bad case of bridesmaid-itis, with no cure in sight. And the gorgeous Dr. Jack Gentry is only making her symptoms worse. Too bad he's already married—to the worst person she knows.

Donna's cute, and she knows it. But with three older sisters who look like young versions of Audrey Hepburn, Andie McDowell, and Isla Fisher—and have effortlessly slim figures to match—cute doesn't quite cut it. Why oh why is Donna the only Gable sister (so far) who takes after their stocky father instead of their slender mother?

Luckily, Donna's larger-than-life personality and her acting talent have always seen her through. But as she watches her beautiful sisters tie the knot, one by one, Donna can't help but wonder when her own leading man will finally take the stage.

To make matters worse, the only guy who has affected Donna's heart rate in years happens to be married to her professional nemesis, Trynn Gentry, who never misses an opportunity to put Donna in her place. There has to be something very wrong with anyone who would enter into a voluntary union with a devious diva like Trynn. So why does Donna's every interaction with Jack feel so very right?